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isopentenyladenine purine alkaloid phytohormone phytohormones growth regulator plant hormone auxin growth rooting cytokinesis cytokinin cytokinin dehydrogenase discadenine synthase metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor cell biology biochemistry biomolecule bioactive molecule molecular formula skeletal structural structure geometry ball stick atom 3d