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homovanillyl alcohol metabolite hydroxytyrosol neurotransmitter dopamine homovanillic mopet 3-methoxy-4-hydroxyphenylethanol 3-methoxy-4-hydroxyphenethyl 4-hydroxy-3-methoxyphenethanol 4-hydroxy-3-methoxyphenethyl phenylethanoid phytochemical antioxidant olives olive leaves oil mediterranean diet novel food 3d receptor agonist neuromodulatory nervous system brain endocannabinoid stress appetite memory signaling neuroscience metabolism biosynthesis mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate synthesis derivative metabolic endogenous