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heme haem holoprotein cytochrome oxygen reduction coordination complex macrocyclic ligand porphyrin chelating iron atom biomolecule bond hemoglobin blood isoprenoid chain metalloprotein stereochemistry cco myoglobin hemeprotein cytochrome oxygen cell biology biochemistry bioactive metabolite biosynthesis mechanism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic endogenous molecule molecular formula skeletal structural structure 3d