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heme haem coordination complex formilporphyrin macrocyclic ligand porphyrin chelating iron atom biomolecule dichroic hemoglobin blood isoprenoid chain metalloprotein cytochrome oxidase stereochemistry cco myoglobin hemeprotein cytochrome oxygen cell biology biochemistry bioactive metabolite biosynthesis mechanism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic endogenous molecule molecular formula skeletal structural structure geometry 3d