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glutamate-1-semialdehyde trna glutamate glutamyl-trna reductase isomerized glutamate-1-semialdehyde 1-aminomutase porphyrins aminolevulinic acid heme chlorophyll rendering metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor human metabolome database hmdb bioactive cell biochemistry biology biomolecule molecule molecular structural structure formula skeletal geometry ball stick tautomer isomer stereoisomer