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gliotoxin sulfur mycotoxin fungi fungus diketopiperazine gliocladium fimbriatum aspergillus fumigatus trichoderma penicillium candida virulence factor immunosuppressive apoptosis immune system poison venom toxicant toxic toxicity toxin toxicology poisoning danger exposure limit intoxication detoxification neutrophils eosinophils granulocytes macrophages thymocytes farnesyl transferase 20s proteasome anti-inflammatory antibiotic antifungal antiviral cytokine cell membrane biosynthesis environmental aspergillosis respiratory immunosuppressed fungal spores immunosuppressive medications 3d