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estradiol palmitate estrogen estrogen ester estradiol monopalmitate hexadecanoate estradiol naturally occurring prohormone receptor affinity estrogenic activity rendering metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor human metabolome database hmdb hormone hormonal therapy replacement function action endocrine endocrinology level analogue