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ergothioneine alkaloid l-ergothioneine thiasine sympectothion ergothionine erythrothioneine thiolhistidinebetaine amino acid histidine derivative fungi actinomycetota cyanobacteria ergot fungus supplement biological activity bioactive nutrient prescription drug food additive nutraceutica dietary liver kidney black beans kidney bean oat bran erythrocytes eye lens semen thiourea imidazoline metabolite metabolism biosynthesis cycle mechanism biosynthetic intermediate biogenesis substrate precursor synthesis bioprecursor metabolic endogenous 3d