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ergosine claviceps purpurea biosynthesis ergotoxine ergoloid-like alkaloid derivative ergot ergotism convulsive gangrenous mycototoxin hallucinogen adrenergic receptor modulator ergolines tryptamine alkaloids quinoline fungi fungus turbina corymbosa ipomoea tricolor convolvulaceae phytotoxin vasoconstriction migraines parkinson's disease oxytocic dopaminergic antidopaminergic serotonergic antiserotonergic lysergic acid amide ergopeptide clavine serotonin biochemistry biomolecule biochemical molecule molecular formula skeletal structural 3d bioprecursor metabolic endogenous