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dihydrotestosterone androstanolone stanolone dht 5a-dihydrotestosterone 5a-dht 3d prostate gland masculine endocrine liver male epididymides metabolism paracrine sex appetite body weight seminal vesicles hormonal masculinization tissues puberty androgen action male genitalia therapy muscular strength hormone masculinizing skeletal muscles backdoor pathway analogue irreversible skin endocrinology biosynthesis embryogenesis sexual differentiation steroid testosterone steroidal diseases benign prostatic hyperplasia