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desmosterol precursor cholesterol bloch pathway biosynthesis dehydrocholesterol reductase desmosterolosis rendering metabolite metabolism cycle mechanism path catabolism biosynthetic intermediate biogenesis substrate synthesis derivative bioprecursor metabolic enzyme endogenous contractor human metabolome database hmdb phosphatidylcholine pc lecithin sphingomyelin phosphatidylethanolamine phospholipid membrane choline cell arachidonic acid lipid phosphocholine fatty ceramide lysophosphatidylcholine phosphatidylserine lipophilic