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crocin carotenoid crocus gardenia flower saffron disaccharide gentiobiose diester antioxidant blood–brain barrier neural protective agent neuroprotective alzheimer's parkinson's disease mood antidepressant antiproliferative anti-cancer anticancer cancer aphrodisiac 3d biosynthetic dietary cell prescription drug monoterpenoid nutrient bioactive coumarin skeletal biological activity catabolism healthy aging plant terpenoid path phytotherapy metabolite essential oils natural molecule mechanism isoflavone biosynthesis herb pathway phytoestrogen