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acetylphosphate acetyl phosphate phosphono acetate dihydrogen acetyl-p acetylphosphoric acid acetylphosphat monoacetyl acetic phosphoric monoanhydride acyl monophosphate escherichia coli acetogenesis reductive acetyl-coa pathway wood-ljungdahl biochemistry metabolite metabolism biosynthesis cycle mechanism path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor molecule molecular formula skeletal structural structure geometry ball stick bioactive cell biomolecule biology 3d