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acetoacetic acid acetoacetate diacetic beta-keto precursor dyes weak biochemistry conjugate base mitochondria liver coenzyme ketone bodies bloodstream fasting type diabetes mellitus succinyl coa citric acid cycle diabetes synthesis diketene tautomerisation keto-enol ketoacidosis ketogenic low-carbohydrate diet nitroprusside ketosis metabolite metabolism biosynthesis mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate derivative bioprecursor metabolic endogenous molecule molecular formula 3d