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dehydrocorticosterone 11-dhc dhc 17-deoxycortisone 11-oxocorticosterone naturally endogenous corticosteroid cortisone corticosterone mineralocorticoid rendering hormone hormonal therapy replacement function action endocrine level prohormone analogue catecholamine norsteroid steroid steroidal metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme contractor