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acceptor adenosine adp atp bilayer chain complex cristae cytochrome cytoplasm dehydrogenase diphosphate donor electrochemical electron energy fad fadh2 gradient harvest harvesting hydrogen illustration intermembrane ion matrix mitochondria mitochondrion nad nadh oxidative oxygen phospholipid phosphorylation plasma power proton quinone redox respiration respiratory scheme transfer transport triphosphate ubiquinol ubiquinone